How can a plant grow fast yet stay stable? How does it sense changes as it grows and reshapes itself? Plant cells are surrounded by a strong cell wall that gives shape, supports growth, and forms materials we use, from wood to fibres. A key problem in agriculture and forestry is a growth–stability trade-off: fast growth often reduces stability, partly because wall properties shift as tissues expand. DYNAMO will test whether chemical and mechanical changes in the wall act as signals that help decide when cells switch from expansion to making stronger tissues. I will use two different plant species —the model plant Arabidopsis thaliana and hybrid aspen— to find these signals and track early responses and signal spread to neighbouring cells. I will then build computational models that link early wall states to later developmental outcomes and test them in hybrid aspen lines. The project will deliver measurable. wall-state descriptors to support earlier selection of crop and tree lines that combine rapid growth with stable structure.